Rose chafer larva starting a back-crawl, note how small are its legs - 2006_06_01

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This unique back-crawl behaviour is a very important indetification clue. All they want to do is to hide from view.
This behaviour was captured on video; see a couple of larvae crawling on paper and crawling on the soil.

Remarkably, these larvae stridulate, not by rubbling their legs like the stag beetle larvae do, but with their mouth parts. They have a maxillo-mandibular stridulation apparatus.

References:

  • Harvey, D.J., Hawes, C.J., Gange, A.C., Finch, P., Chesmore, D. & Farr, I. (2011)  Development of non-invasive monitoring methods for larvae and adults of the stag beetle, Lucanus cervus. Insect Conservation and Diversity 4: 4-14. DOI: 10.1111/j.1752-4598.2009.00072.x [PDF]
  • Wessel, A. (2006)   Stridulation in the Coleoptera - An Overview. In: Drosopoulos, S. & Claridge, M.F. (eds). Insect Sounds and Communication. Physiology, Behaviour, Ecology and Evolution. (Contemporary Topics in Entomology) CRC-Taylor & Francis, Boca Raton, London, New York: 397-403. [PDF]